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Masaki Miya, Matt Friedman, Takashi P. Satoh, Hirohiko Takeshima, Tetsuya Sado, Wataru Iwasaki, Yusuke Yamanoue, Masanori Nakatani, Kohji Mabuchi, Jun G. Inoue, Jan Yde Poulsen, Tsukasa Fukunaga, Yukuto Sato, and Mutsumi Nishida. Evolutionary origin of the Scombridae (tunas and mackerels): Members of a paleogene adaptive radiation with 14 other pelagic fish families. PLOS ONE, 8, e73535. (2013)